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The orange car you were refering to with the new lights and the stock taillights just might have been the first car completed in late 69. Perhaps Joel was not happy with the stock look of the rear, so he sent it back to have the taillights re-done. That car just may be the one with the Dodge Charger pop-up gascap, unlike all the rest with the "MOON" popup cap.
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I do think that was the first customer car built but I don't think it became the car with the Dodge gas cap. I think it's an unknown/unfound car today... Or maybe someone out there knows exactly where/what it is today..
Here are a few comparitive pics..
Here are 2 of the orange/black GT with stock taillights/open headlights:
Note this one has the daytona/moon gas cap..