As to my comment above, I forgot about the black backside. Thanks for correcting me.
As to the body colored wheels, I'm only aware of 6 cars with them. One 1967 and five 68's. No 69's.
4 of those 68 cars were special paint, built in April or May. For non-rally wheel cars, the wheel color matches the body color. It's not a big leap that the special order color was inadvertently applied to the wheels, even though they were rally wheels. I mean who would think to order body color rallys?
The 67 car had a broadcast sheet that survives. Jon, are there any pictures of that 67 with the original wheels?
That leaves the other 68. October build, just a regular paint code on the trim tag and POP. Dunno.